The direct revenue subscriptions sales is the most straight forward way to monetize. Split pricing options for whales to maximise the revenue potential from your biggest fans.
But revenue models can (should) be combined. E.g. Commercial partnerships and sponsorship deals are the most common way to valorise the user base to other players.
Many more techniques can be explored: upselling sister-brands, shared logistic/distribution services, data sales & new insights from observed consumer behavior.